| Serovar | Strain | Serogroup | Country | Source |
|---|---|---|---|---|
| sofia | Sofia 874 | 12 | Bulgaria | Human |
| perameles | Bandicoot 343 | 16 | Australia | Perameles nasuta |
| ranarum | Iowa City Frgo (ICF) | 22 | USA | Frog |
| hardjo | Went 5 | 22 | Canada | ? |
| semaranga | Veldrat Semarang 173 | 28 | Indonesia | Rat |
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